We gratefully acknowledge Laura Cutire, Laura Ramirez, Javier F. Valles, and Erika Sajami for the field work. We also thank the staff of Alwyn Gentry Herbarium of Universidad Nacional Amaz?nica de Madre de Dios and of San Marcos Herbarium of Universidad Nacional Mayor de San Marcos for their technical support in the taxonomic identification. We also thank FONDECYT-CONCYTEC (Fondo Nacional de Desarrollo Cient?fico, Tecnol?gico y de Innovaci?n Tecnol?gica, Per?, award #239-2018) for the fellowship grant to RC-L; CAPES (Coordena??o de Aperfei?oamento de Pessoal de N?vel Superior do Governo do Brasil, Finance Code 001 and #88882.305844/2018-01), FAPESP (Funda??o de Amparo ? Pesquisa do Estado de S?o Paulo, grant #2012/24118-8), and The Royal Society for the fellowship granted to DR; and also FAPESP (grant #2018/22914-8) for the fellowship grant to DOR. Funding. Funding for the field work was...