DGS and JAB were funded by a Sir Henry Dale Fellowship, jointly funded by the Wellcome Trust and Royal Society [Grant 102507/Z/13/Z]. DGS was funded by a Wellcome Trust Senior Research Fellowship [217221/Z/19/Z]. CS, JAB, NF and DGS were also funded by a CONCYTEC-UK Embassy grant [No. 003-2016-FONDECYT]. JAB was funded by the National Fund for Scientific and Technological development of Chile [FONDECYT-Iniciación, grant number 11181017]. DF-C receives a doctoral fellowship from the National Agency for Research and Development (ANID) [process number BCH 72170436]. NL is a research fellow of CNPq [312249/2017-9].